He says camera traps have identified 24 and the rest have been calculated using the unreliable pugmark method.
据他说,照相机拍摄到24只,而其余的则是依据足迹等不太可靠的方法计算得来。
The researchers then use the pictures to measure the length and width of each pugmark, the angle between toes, and 10 other variables to establish an ID.